22:duplication&14:speciation&10:speciation&2:duplication&0:leaf&1:leaf&9:creation&7:creation&5:creation&3:leaf&4:leaf&6:leaf&8:leaf&13:speciation&11:leaf&12:leaf&21:speciation&17:speciation&15:leaf&16:leaf&20:speciation&18:leaf&19:leaf